(1997 ) reported elymus coreanus ( h . coreand ) and e . californicus ( h . californicd ) have the same genome constitution of leymus ( nsxm ), and included them in leymus coreana)和ecalifornicus(hcalifornica)具有與賴草屬leymus相同的nsxm染色體組,并把這兩個物種組合到leymus中,使得hystrix的分類更為復(fù)雜。